User blog comment:IamJakuhoRaikoben/Chapter 444: Emperor Spriggan, Review./@comment-5288632-20150713060134

And so, Mest teleported them to an underwater area, only to be vehicle controlled by Sorano, A.K.A Angel. Sure, she has to remind Lucy that she's the one who killed Karen in front of her face, but then again, Karen is still a despicable person for abusing Aries. We were finally able to see Makarov at the one year time skip. And, boy, does he look different thanks to the art evolution. And on a surprising note, the emperor of the Spriggan Empire is none other than...Zeref! I guess that means that God Serana decided to become Zeref's right-hand man...or is it? I wonder what happened to Acnologia? One thing for sure is that dragon won't go down to easily. Anyway, Synchronicity time! :)